The story takes place in 2018, after Judgment Day, when "Skynet," ruled by autonomously awakened AIs, launched a nuclear attack on humanity. John Connor, who has become the leader of the Resistance, takes command on the front lines of the battle against the Machine Army, which is further strengthening its equipment to eliminate all humans. In a time loop setting, Marcus Wright works with Kyle Reese, who becomes John Connor's father in the near future. He was once a powerful cyborg created by fusing machines and living organisms as research material for Skynet. The Harvester is a giant mecha that harvests the human survivors alive on the ground. Marcus Wright, having lost the memory of his own creation, is now fighting Kyle Reese and the giant Harvester. The mecha that separates from the Harvester leg and relentlessly pursues it is the Moto Terminator.